1. Department of Evolution, Ecology and Behavior, University of Illinois Urbana–Champaign, Urbana, IL 61801, USA
2. Carl R. Woese Institute for Genomic Biology, University of Illinois Urbana–Champaign, Urbana, IL 61801, USA
3. Program in Ecology, Evolution and Conservation, University of Illinois Urbana–Champaign, Urbana, IL 61801, USA