1. Wellcome Trust Centre for Cell‐Matrix Research, Faculty of Biology, Medicine and HealthUniversity of ManchesterOxford RoadManchesterM13 9PT United Kingdom
2. Department of BioengineeringUniversity of PittsburghPittsburgh Pennsylvania15213
3. Department of Developmental BiologyUniversity of PittsburghPittsburgh Pennsylvania15213
4. Department of Computational and Systems BiologyUniversity of PittsburghPittsburgh Pennsylvania15213