1. Frankincense and Biodiversity Group, Dhofar University, Salalah 2509, Oman
2. School of Environment and Science, Griffith University, 170 Kessels Rd, Nathan, Queensland 4111, Australia
3. School of Biology, Ecole de Biologie Industrielle (EBI), Cergy, France
4. Oman Pharmaceutical Products Co. LLC, Salalah, Oman
5. Environmental Futures Research Institute, Nathan Campus, Griffith University, 170 Kessels Rd, Nathan, Queensland 4111, Australia