1. State Key Laboratory of Desert and Oasis Ecology, Xinjiang Institute of Ecology and Geography, Chinese Academy of Sciences, Urumqi, 830011, PR China
2. University of the Chinese Academy of Sciences, Beijing 100049, PR China
3. State Key Laboratory of Microbial Resources and Environmental Microbiology Research Center at Institute of Microbiology, Chinese Academy of Sciences, Beijing 100101, PR China
4. State Key Laboratory of Microbial Biotechnology, Shandong University, Qingdao 266237, PR China