1. Soonchunhyang Institute of Medi-bio Science, Soonchunhyang University 1 , Cheonan 31151 , Korea
2. Soonchunhyang University 2 Department of Integrated Biomedical Science , , Cheonan 31151 , Korea
3. Center for RNA Research, Institute for Basic Science 3 , Seoul 08826 , Korea
4. School of Biological Sciences, Seoul National University 4 , Seoul 08826 , Korea