1. These authors contributed equally to this work
2. Influenza Virus Research Center, National Institute of Infectious Diseases, Tokyo, Japan
3. Division of Virology, Department of Microbiology and Immunology, Institute of Medical Science, University of Tokyo, Tokyo, Japan
4. Influenza and Prion Disease Research Center, National Institute of Animal Health, National Agriculture and Food Research Organization, Ibaraki, Japan
5. Department of Pathobiological Sciences, School of Veterinary Medicine, University of Wisconsin-Madison, Madison, United States
6. The United Graduate School of Veterinary Sciences, Gifu University, Gifu, Japan
7. Department of Special Pathogens, International Research Center for Infectious Diseases, Institute of Medical Science, University of Tokyo, Tokyo, Japan
8. ERATO Infection-Induced Host Responses Project, Japan Science and Technology Agency, Saitama, Japan