1. Department of Biology and Centre for Forest Biology, University of Victoria, Victoria, British Columbia, Canada V8W 3N5 (M.Z., A.G., C.P.C.); Department of Agricultural, Food, and Nutritional Science, University of Alberta, Edmonton, Alberta, Canada T6G 2P5 (A.J., J.A.O.); Plant Biotechnology Institute, National Research Council of Canada, Saskatoon, Saskatchewan, Canada S7N 0W9 (L.I.Z., S.R.A.);