Abstract
The notion of well-being is relative to the needs of organisms, both non-human and human. I argue that what we have learned about development, animal consciousness and ecological interrelations requires an evolution-informed extension of the way we think about well- and ill-being. Studies of epigenetic inheritance, mostly in animals, but also in humans, show that the effects of both stress and well-being go deeper ‘under the skin’ and extend further into the future than we thought. Studies of animals’ consciousness suggest that welfare considerations should be extended to many more animal species than most of us imagined. Technological innovations and ecological studies show that well-being depends critically, far more than we anticipated, on global social and environmental politics, social justice and the uses of technology.