1. Department of Agricultural Biotechnology and Research Institute of Agriculture and Life Sciences, Seoul National University, Seoul 08826, Republic of Korea
2. Institute of Agricultural Science and Technology, Kyungpook National University, Daegu 41566, Republic of Korea
3. School of Food Science and Biotechnology, Kyungpook National University, Daegu 41566, Republic of Korea
4. Major of Beef Science, Department of Livestock, Korea National University of Agriculture and Fisheries, Jeonju 54874, Republic of Korea
5. Division of Animal and Dairy Science, Chungnam National University, Daejeon 34134, Republic of Korea