1. Department of Biology, Kyung Hee University , 26 Kyungheedae-ro, Dongdaemun-gu, Seoul 02447 , Republic of Korea
2. Department of Biological Science, College of Bioscience and Biotechnology, Chungnam National University , Daejeon 34134 , Republic of Korea
3. National Herbarium of New South Wales, Australian Institute of Botanical Science, Royal Botanic Gardens & Domain Trust , Locked Bag 6002, Mount Annan, NSW 2567 , Australia