1. Department of Biology, Faculty of Science, Shinshu University , Asahi 3-1-1, Matsumoto, Nagano, 390-8621 Japan
2. Research Institute for Sustainable Humanosphere, Kyoto University , Gokasho, Uji, Kyoto, 611-0011 Japan
3. Department of Applied Biology, Faculty of Life Sciences, Toyo University , Izumino 1-1-1, Itakura-machi, Oru-gun, Gunma, 374-0193 Japan