1. State Key Laboratory of Protein and Plant Gene Research, School of Advanced Agricultural Sciences, Peking University , Beijing 100871 , China
2. Guangdong Laboratory for Lingnan Modern Agriculture, College of Life Sciences, South China Agricultural University , Guangzhou 510642 , China
3. School of Life Sciences, Peking University , Beijing 100871 , China
4. State Key Laboratory of Genetic Engineering, Institute of Plant Biology, School of Life Sciences, Fudan University , Shanghai 200438 , China
5. Shandong Laboratory of Advanced Agricultural Sciences at Weifang , Peking University Institute of Advanced Agricultural Sciences, Shandong 261000 , China