1. Department of Biology, Christian Brothers University, Memphis, TN 38104, USA
2. Edward J. Meeman Biological Field Station, The University of Memphis, Millington, TN 38058, USA
3. School of Biological Sciences, The University of Northern Colorado, Greeley, CO 80639, USA
4. Highlands-Cashiers Land Trust, Highlands, NC 28723, USA
5. Department of Biology, Western Carolina University, Cullowhee, NC 28723, USA