1. Department of Biology and Environmental Sciences, Carl von Ossietzky University of Oldenburg, Carl-von-Ossietzky-Straße 9-11, D-26111 Oldenburg, Germany
2. Department of Microbiology and Ecosystem Science, University of Vienna, Althanstrasse 14, A-1090 Vienna, Austria
3. Smithsonian Tropical Research Institute, Apartado Postal 0843-03092, Balboa, Ancon, Panamá, República de Panamá