1. Guangdong Provincial Key Laboratory of Applied Botany, South China Botanical Garden, Chinese Academy of Sciences , Guangzhou 510650 , China
2. Southern Marine Science and Engineering Guangdong Laboratory (Guangzhou) , Guangzhou 511458 , China
3. Key Laboratory of Vegetation Restoration and Management of Degraded Ecosystems, Key Laboratory of Carbon Sequestration in Terrestrial Ecosystem, South China Botanical Garden, Chinese Academy of Sciences , Guangzhou 510650 , China
4. Bioversity International, Parc Scientifique Agropolis II , 34397 Montpellier , France
5. CIRAD, UMR AGAP Institut , F-34398 Montpellier , France
6. UMR AGAP Institut, Univ Montpellier, CIRAD, INRAE, Institut Agro , Montpellier , France
7. Key Laboratory of Plant Resources Conservation and Sustainable Utilization, South China Botanical Garden, Chinese Academy of Sciences , Guangzhou 510650 , China
8. Department of Genetics and Genome Biology, University of Leicester , Leicester LE1 7RH , UK