1. Department of Earth , Ocean and Atmospheric Sciences, , 2039–2207 Main Mall, Vancouver, BC V6T 1Z4 , Canada
2. University of British Columbia , Ocean and Atmospheric Sciences, , 2039–2207 Main Mall, Vancouver, BC V6T 1Z4 , Canada
3. Institute for the Oceans and Fisheries, University of British Columbia , 2202 Main Mall, Vancouver, BC V6T 1Z4 , Canada
4. Scripps Institution of Oceanography, University of California , 9500 Gilman Drive, La Jolla, CA 92093 , USA
5. Virginia Institute of Marine Science, William & Mary , 1370 Greate Road, Gloucester Point, VA 23062 , USA