1. Boyce Thompson Institute , Ithaca, New York 14853, USA
2. Plant Biology Section, School of Integrative Plant Science, Cornell University , Ithaca, New York 14853, USA
3. Plant Pathology and Plant-Microbe Biology Section, School of Integrative Plant Science, Cornell University , Ithaca, New York 14853, USA
4. U.S. Department of Agriculture-Agricultural Research Service, Robert W. Holley Center for Agriculture and Health , Ithaca, New York 14853, USA