1. Division of Biology, Kansas State University , Manhattan, Kansas 66506, USA
2. Molecular, Cellular and Developmental Biology, Kansas State University , Manhattan, Kansas 66506, USA
3. Donald Danforth Plant Science Center , Olivette, Missouri 63132, USA
4. Department of Botany and Plant Sciences, University of California , Riverside, California 92521, USA
5. Interdepartmental Genetics, Kansas State University , Manhattan, Kansas 66506, USA
6. Department of Molecular and Integrative Physiology, University of Kansas Medical Center , Kansas City, Kansas 66160, USA