1. Cilia Cell Biology, Institute of Molecular Physiology, Johannes Gutenberg University Mainz , 55128 Mainz , Germany
2. Imaging Core Facility, Cell Biology Unit, University Medical Centre, Johannes Gutenberg University Mainz , 55101 Mainz , Germany
3. Randall Centre for Cell and Molecular Biophysics, King's College London , London SE1 1UL , UK