1. Department of Natural History Sciences, Graduate School of Science, Hokkaido University , Sapporo, Hokkaido 060-0810, Japan
2. Department of Oceanography, University of Hawaii at Manoa , Honolulu, HI 96822, United States
3. Institute for the Advancement of Higher Education, Hokkaido University , Sapporo, Hokkaido 060-0817, Japan
4. Graduate School of Science, Hokkaido University , Sapporo, Hokkaido 060-0810, Japan