1. Department of Microbiology and Immunology, Rega Institute, KU Leuven, Leuven, Belgium
2. Institute of Evolutionary Biology, University of Edinburgh, Edinburgh, UK
3. Centre for Immunology, Infection and Evolution, University of Edinburgh, Ashworth Laboratories, King’s Buildings, Edinburgh, UK
4. Department of Human Genetics, David Geffen School of Medicine, University of California, Los Angeles, CA, USA
5. Department of Biostatistics, School of Public Health, University of California, Los Angeles, CA, USA
6. Department of Biomathematics, David Geffen School of Medicine, University of California, Los Angeles, CA, USA