1. School of Science, Mae Fah Luang University , Chiang Rai 57100 , Thailand
2. Division of Biological Sciences, College of Arts and Sciences, University of the Philippines Visayas , Miagao, Iloilo 5024 , Philippines
3. School of Life Sciences, University of Warwick , Gibbet Hill Road, Coventry CV4 7AL , United Kingdom
4. Department of Biosciences, Swansea University , Swansea SA2 8PP , United Kingdom