1. Division of Ecology and Evolutionary Biology, Graduate School of Life Sciences, Tohoku University, Sendai, Miyagi, Japan
2. Department of Biology, Faculty of Science, Yamagata University, Yamagata, Yamagata, Japan
3. Ohba Firefly Research Laboratory, Yokosuka, Kanagawa, Japan
4. Research Institute for Cell Engineering, National Institute of Advanced Industrial Science and Technology, Osaka, Japan