1. Department of Advanced Bioscience, Graduate School of Agriculture, Kindai University , Nara City , Nara 631-8505 , Japan
2. Laboratory of Genome and Chromosome Functions, Institute for Protein Research, Osaka University , 3-2 Yamadaoka, Suita , Osaka 565-0871 , Japan
3. Agricultural Technology and Innovation Research Institute, Kindai University , Nara City , Nara 631-8505 , Japan