1. Laboratory of Evolutionary Genetics, Institute of Biology, University of Neuchâtel , Neuchâtel , Neuchâtel 2000 , Switzerland
2. Department of Plant Pathology, University of Wisconsin-Madison , Madison , WI 53706 , USA
3. Wisconsin Institute for Discovery , Madison , WI 53706 , USA
4. Systematic Biology, Department of Organismal Biology, Uppsala University , Uppsala , 752 36 , Sweden