1. Department of Clinical and Biomedical Sciences, University of Exeter , 79 Heavitree Road, Exeter, EX2 4TH , United Kingdom
2. Exeter Genomics Laboratory , RILD Building, , Barrack Road, Exeter, EX2 5DW , United Kingdom
3. Royal Devon University Healthcare NHS Foundation Trust , RILD Building, , Barrack Road, Exeter, EX2 5DW , United Kingdom
4. Wellcome Centre for Mitochondrial Research , Translational and Clinical Research Institute, , Framlington Place, Newcastle upon Tyne, NE2 4HH , United Kingdom
5. Newcastle University , Translational and Clinical Research Institute, , Framlington Place, Newcastle upon Tyne, NE2 4HH , United Kingdom