1. Curriculum in Genetics and Molecular Biology, The University of North Carolina at Chapel Hill , Chapel Hill, NC 27599 , USA
2. Department of Biology, The University of North Carolina at Chapel Hill , Chapel Hill, NC 27599 , USA
3. Department of Genetics, The University of North Carolina at Chapel Hill , Chapel Hill, NC 27599 , USA
4. Integrative Program for Biological and Genome Sciences, The University of North Carolina at Chapel Hill , Chapel Hill, NC 27599 , USA