1. Department of Biochemistry and Molecular Biology, Saitama University , Saitama, 338-8570 Japan
2. Suntory Rising Stars Encouragement Program in Life Sciences (SunRiSE), Suntory Foundation for Life Sciences , Kyoto, 619-0284 Japan
3. Department of Botany, University of Wisconsin , Madison, WI 53706, USA
4. College of Plant Science and Technology, Huazhong Agricultural University , Wuhan 430070, China