1. Afromontane Research Unit & Department of Zoology & Entomology, University of the Free State , Private Bag X13, Phuthaditjhaba 9866 , South Africa
2. School of Mathematical & Natural Science, University of Venda , Thohoyandou, 0950 , South Africa
3. Wild Bird Trust, National Geographic Okavango Wilderness Project , 27 Wolfridge Road, Hogsback 5721 , South Africa
4. Centre for Viral Zoonoses, Department of Medical Virology, University of Pretoria , Pretoria 0002 , South Africa
5. Durban Natural Science Museum , P.O. Box 4085, Durban 4000 , South Africa
6. School of Mathematical & Natural Science, University of Venda , Thohoyandou , 0950, South Africa
7. Ditsong Natural History Museum , Pretoria 0002 , South Africa
8. Department of Zoology and Entomology, University of Pretoria , Private Bag 20, Hatfield 0028, Pretoria , South Africa
9. Geoecodynamics Research Hub, c/o Department of Botany and Zoology, University of Stellenbosch , Private Bag X1 Matieland, 7602, Stellenbosch , South Africa
10. Mammal Research Institute, Department of Zoology and Entomology, University of Pretoria, Private Bag 20, Hatfield 0028, Pretoria , South Africa
11. Department of Biological Sciences, University of Eswatini , Private Bag 4, Kwaluseni , Eswatini