1. College of Fisheries, Key Lab of Freshwater Animal Breeding, Ministry of Agriculture and Rural Affairs/Key Lab of Agricultural Animal Genetics, Breeding and Reproduction of Ministry of Education/Engineering Research Center of Green Development for Conventional Aquatic Biological Industry in the Yangtze River Economic Belt, Ministry of Education, Huazhong Agricultural University , Wuhan 430070 , China
2. Hubei Hongshan Laboratory , Wuhan 430070 , China
3. CAS Key Laboratory of Tropical Marine Bio-Resources and Ecology, South China Sea Institute of Oceanology, Innovation Academy of South China Sea Ecology and Environmental Engineering, Chinese Academy of Sciences , Guangzhou 510301 , China
4. Department of Biology, Ghent University , Ghent B-9000 , Belgium
5. Unit of Biotechnology, University of Mogi das Cruzes , Mogi das Cruzes, Săo Paulo 08780-911, Brazil
6. Guangdong Laboratory for Lingnan Modern Agriculture , Guangzhou 510000 , China