1. Edinburgh Neuroscience, School of Biomedical Sciences, The University of Edinburgh , Edinburgh EH8 9JZ , UK
2. Family Centre for Research with Ubuntu (FAMCRU), Tygerberg Hospital, Department of Paediatrics and Child Health, Stellenbosch University , Cape Town 7505 , South Africa
3. Department of Global Health and Infection, Brighton and Sussex Medical School, University of Sussex , Brighton BN2 5BE , UK