1. Department of Bionanoscience, Kavli Institute of Nanoscience Delft, Delft University of Technology , Delft , the Netherlands
2. Department of Fundamental Microbiology, Faculty of Biology and Medicine, University of Lausanne , Lausanne , Switzerland
3. School of Physics and Astronomy, University of Edinburgh , Edinburgh , UK
4. MRC Human Genetics Unit, Institute of Genetics and Cancer, University of Edinburgh , Edinburgh , UK