1. Department of Developmental Biology, Stanford University School of Medicine , Stanford, CA , USA
2. State Key Laboratory of Non-food Biomass and Enzyme Technology, Guangxi Academy of Sciences , 530007 Nanning , China
3. IBS Center for Genomic Integrity and Department for Biological Sciences, Ulsan National Institute of Science and Technology , Ulsan , Korea
4. Swiss Institute for Experimental Cancer Research (ISREC) and School of Life Sciences, Swiss Federal Institute of Technology Lausanne (EPFL) , Lausanne , Switzerland
5. Department of Genetics, Stanford University School of Medicine , Stanford, CA , USA