1. CSIC, Global Ecology Unit, CREAF-CSIC-UAB, 08193, Cerdanyola del Vallès, Catalonia, Spain
2. CREAF, 08193, Cerdanyola del Vallès, Catalonia, Spain
3. Department of Ecology and Genetics, P.O. Box 3000, FI-90014, University of Oulu, Oulu, Finland
4. Institute for Research in Biodiversity (IRBio) and Department of Evolutionary Biology, Ecology and Environmental Sciences, University of Barcelona, Av. Diagonal 643, 08028, Barcelona, Catalonia, Spain
5. Naturalis Biodiversity Center, Vondellaan 55, P.O. Box 9517, Leiden, The Netherlands
6. Faculty of Science, Leiden University, P.O. Box 9502, 2300 RA Leiden, The Netherlands