1. Department of Ecology, Environment and Evolution, La Trobe University, Melbourne, Victoria, Australia
2. Kings Park Science, Department of Biodiversity Conservation and Attractions, Kings Park, WA, Australia
3. Ecology and Evolution, Research School of Biology, The Australian National University, Canberra, ACT, Australia
4. Royal Botanic Gardens Victoria, Corner of Ballarto Road and Botanic Drive, Cranbourne, VIC, Australia