1. Department of Life Sciences , Graduate School of Arts and Sciences, , 3-8-1 Komaba, Meguro, Tokyo 153-8902 , Japan
2. The University of Tokyo , Graduate School of Arts and Sciences, , 3-8-1 Komaba, Meguro, Tokyo 153-8902 , Japan
3. Department of Neuropsychiatry, Keio University School of Medicine , 35 Shinanomachi, Shinjuku, Tokyo 160-8582 , Japan
4. Teijin Pharma Limited , 4-3-2 Asahigaoka, Hino, Tokyo 191-8512 , Japan