1. Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming, Yunnan, China
2. College of Life Sciences, Guizhou University, Guiyang, Guizhou, China
3. Museum of Nature and Human Activities, Hyogo, Yayoigaoka, Sanda, Hyogo, Japan
4. Department of Biology, University of Nebraska-Kearney, Kearney, NE, USA
5. Department of Biology, University of Florida, Gainesville, FL, USA
6. Florida Museum of Natural History, University of Florida, Gainesville, FL, USA