1. The Hakubi Center for Advanced Research, Kyoto University, Yoshida-Konoe-cho, Kyoto, Japan
2. Seto Marine Biological Laboratory, Field Science, Education and Research Center, Kyoto University, Wakayama, Japan
3. Department of Gene Mechanisms, Graduate School of Biostudies, Kyoto University, Yoshida-Konoe-cho, Kyoto, Japan