1. Kennedy Institute of Rheumatology, University of Oxford, Oxford, UK
2. Translational Gastroenterology Unit, Nuffield Department of Clinical Medicine, University of Oxford, Oxford, UK
3. MRC Human Immunology Unit, Weatherall Institute of Molecular Medicine, Radcliffe Department of Medicine, University of Oxford, Oxford, UK
4. Wellcome Trust Centre for Human Genetics, Radcliffe Department of Medicine, University of Oxford, Oxford, UK
5. Sir William Dunn School of Pathology, University of Oxford, Oxford, UK