1. Department of Ecology and Evolutionary Biology, Yale University, New Haven, Connecticut 06520
2. Microbial Sciences Institute, Yale University, West Haven, Connecticut 06516
3. Yale Institute for Biospheric Studies, Yale University, New Haven, Connecticut 06520
4. Department of Global Ecology, Carnegie Institution for Science, Stanford, California 94305
5. Department of Ecology and Evolutionary Biology, University of California, Irvine, California 92697