1. School of Biological and Marine Sciences, University of Plymouth, Plymouth, Devon, UK
2. Centre for Ecology & Conservation, University of Exeter, Penryn, Cornwall, UK
3. Department of Integrative Biology, University of Guelph, Guelph, Ontario, Canada
4. Department of Research, Área de Conservación Guanacaste, Liberia, Guanacaste, Costa Rica
5. School of Biology, Universidad de Costa Rica, San Pedro, San Jose, Costa Rica
6. Department of Biology, University of Pennsylvania, Philadelphia, PA, United States of America
7. Guanacaste Dry Forest Conservation Fund, Philadelphia, United States of America