1. Paul F. Glenn Center for Biology of Aging Research, Department of Genetics, Blavatnik Institute, Harvard Medical School (HMS), Boston, MA 02115, USA
2. Molecular and Biomedical Sciences, University of Maine, Orono, ME 04467, USA
3. Division of Genetics, Department of Medicine, Brigham and Women’s Hospital, Harvard Medical School, Boston, MA 02115, USA
4. Belozersky Institute of Physico-Chemical Biology, Moscow State University, Moscow 119234, Russia
5. Department of Biology and Chemistry, Massachusetts Institute of Technology, Cambridge, MA 02142, USA