1. Key Laboratory of Animal Ecology and Conservation Biology, Institute of Zoology, Chinese Academy of Sciences, Beijing, China
2. Key Laboratory of Adaptation and Evolution of Plateau Biota, Northwest Institute of Plateau Biology, Chinese Academy of Sciences, Xining, Qinghai, China
3. Xining Wildlife Zoo, Xining, Qinghai, China
4. Beijing Zoo, Beijing, China
5. Chongqing Zoo, Chongqing, China