1. Fujian Agriculture and Forestry University, Fuzhou, China
2. Key Laboratory of Animal Pathogen Infection and Immunology of Fujian Province, College of Animal Sciences, Fujian Agriculture and Forestry University, Fuzhou, China
3. CAS Key Laboratory of Pathogenic Microbiology and Immunology, Institute of Microbiology, Chinese Academy of Sciences (CAS), Beijing, China
4. Department of Virology, Faculty of Veterinary Medicine, Suez Canal University, Egypt