1. School of Molecular Biosciences, College of Veterinary Medicine, Washington State University , Pullman, Washington, USA
2. RNA Viruses Section, Laboratory of Infectious Diseases, National Institute of Allergy and Infectious Diseases, National Institutes of Health , Bethesda, Maryland, USA
3. Paul G. Allen School for Global Health, College of Veterinary Medicine, Washington State University , Pullman, Washington, USA