1. Laboratory of Systems Virology, Institute for Frontier Life and Medical Sciences, Kyoto University, Kyoto, Japan
2. Graduate School of Pharmaceutical Sciences, Kyoto University, Kyoto, Japan
3. Division of Systems Virology, Department of Infectious Disease Control, International Research Center for Infectious Diseases, Institute of Medical Science, the University of Tokyo, Tokyo, Japan
4. Graduate School of Medicine, the University of Tokyo, Tokyo, Japan
5. Department of Molecular Virology, Tokyo Medical and Dental University, Tokyo, Japan
6. Graduate School of Medicine, Kyoto University, Kyoto, Japan
7. Graduate School of Biostudies, Kyoto University, Kyoto, Japan
8. MRC-University of Glasgow Centre for Virus Research, University of Glasgow, Glasgow, United Kingdom
9. Department of Animal Science, Faculty of Agriculture, Tokyo University of Agriculture, Kanagawa, Japan
10. Department of Genomic Function and Diversity, Medical Research Institute, Tokyo Medical and Dental University, Tokyo, Japan
11. CREST, Japan Science and Technology Agency, Saitama, Japan