1. Graduate School of Agriculture, Tokyo University of Agriculture and Technology, Fuchu, Tokyo, Japan
2. International Center for Isotope Effects Research, School of Earth Sciences and Engineering, Nanjing University, Nanjing, Jiangsu, China
3. Faculty of Life and Environmental Sciences, University of Tsukuba, Tsukuba, Ibaraki, Japan
4. Euglena Co., Ltd., Minato‑ku, Tokyo, Japan
5. Department of Biology, Clark University, Worcester, Massachusetts, USA
6. Independent Administrative Institution, Tokyo National Research Institute for Cultural Properties, Taito-ku, Tokyo, Japan