1. School of Biology, Faculty of Biological Sciences, University of Leeds, Leeds, United Kingdom
2. School of Biomedical Sciences, Faculty of Biological Sciences, University of Leeds, Leeds, United Kingdom
3. Institute of Immunity & Transplantation, UCL Department of Virology, Division of Infection & Immunity, Royal Free Hospital, London, United Kingdom
4. School of Molecular and Cellular Biology, Faculty of Biological Sciences, University of Leeds, Leeds, United Kingdom
5. Faculty of Natural Sciences, University of Keele, Newcastle-under-Lyme, United Kingdom