1. a Department of Biological Sciences, Korea Advanced Institute of Science and Technology, Daejeon 305-701, South Korea
2. b Department of Biological Sciences, Korea Advanced Institute of Science and Technology, Daejeon 305-701, South Korea
3. c Department of Biological Sciences, Korea Advanced Institute of Science and Technology, Daejeon 305-701, South Korea
4. d Department of Biological Sciences, Korea Advanced Institute of Science and Technology, Daejeon 305-701, South Korea
5. e Department of Biological Sciences, Korea Advanced Institute of Science and Technology, Daejeon 305-701, South Korea