1. Department of Applied Biology, Kangwon National University, Chuncheon 24341, Korea
2. Department of Horticultural Biotechnology, College of Life Sciences, Kyung Hee University, Yongin 17104, Korea
3. Animal and Plant Quarantine Agency, Gimcheon 39660, Korea
4. Crop Protection, National Academy of Agricultural Science, Rural Development Administration, 166, Nongsaengmyeong-ro, Iseo-myeon, Wanju-gun, Jeollabuk-do 55365, Korea
5. Department of Horticultural Biotechnology, College of Life Sciences, Kyung Hee University, Yongin 17104, Korea. This work was supported by a grant from the Animal and Plant Quarantine Agency (2013-2015), Korea, and the Cooperative Research Program for Agriculture Science & Technology Development (Project No. PJ011758022016), Rural Development Administration, Republic of Korea.